Submitted OR
In review

Dimensions of Marine Phytoplankton Diversity. S. Dutkiewicz, P. Cermeno, O. Jahn, M.J. Follows, A. Hickman, D. Taniguchi and B.A. Ward. Biogeosciences Discussions.

Nutrients, ecosystems, and the expansion of eukaryotic life. C.T. Reinhard, N.J. Planavsky, B.A. Ward, G.D. Love, G. Le Hir and A.J. Ridgwell.


Considering the role of adaptive evolution in models of the ocean and climate system. B.A. Ward, S. Collins, S. Dutkiewicz, S. Gibbs, P. Bown, A.J. Ridgwell, B. Sauterey, J.D. Wilson and A. Oschlies. [Invited contribution to the AGU100 “Grand Challenges in Earth and Space Science”.]


Mixotroph ecology: more than the sum of its parts. B.A. Ward (2019). Proceedings of the National Academy of Sciences, USA, 116 (13) 5846-5848. [Invited Commentary on K. Edwards (2019): Mixotrophy in nanoflagellates across environmental gradients in the ocean. Proceedings of the National Academy of Sciences, USA, 116 (13) 6211-6220.]

A trait-based modelling approach to planktonic foraminifera ecology. M. Grigoratou, F. M. Monteiro, D. N. Schmidt, J. D. Wilson, B. A. Ward and A. J. Ridgwell. Biogeosciences, 16, 1469-1492.


EcoGEnIE 0.1: Plankton Ecology in the cGENIE Earth system model. B.A. Ward, J.D. Wilson, R.M. Death, F.M. Monteiro, A. Yool and A.J. Ridgwell (2018). Geoscientific Model Development,

Linking marine plankton ecosystems and climate: A new modelling approach to the warm Early Eocene climate. J.D. Wilson, F.M. Monteiro, D.N. Schmidt, B.A. Ward and A.J. Ridgwell (2018). Paleoceanography and Paleoclimatology, 33 (12) 1439-1452.

Theoretical interpretations of subtropical plankton biogeography. M.J. Follows, S. Dutkiewicz, B.A. Ward and C. Follet (2018). In Gasol, Josep and Kirchman, David (eds.) Microbial Ecology of the Oceans. 3rd ed. Hoboken, NJ. John Wiley, p. 467.


The Size Dependence of Phytoplankton Growth Rates: A Trade-Off between Nutrient Uptake and Metabolism. B.A. Ward, E. Marañón, B. Sauterey, J. Rault and D. Claessen (2017). The American Naturalist, 189 (2) 170-177.

The implications of eco-evolutionary processes for the emergence of marine plankton community biogeography. B. Sauterey, B.A. Ward, J. Rault, C. Bowler and D. Claessen (2017). The American Naturalist, 190 (1) 116-130.

Assessing an efficient “Instant Acclimation” approximation of dynamic phytoplankton stoichiometry. B.A. Ward (2017). Journal of Plankton Research, 39 (5) 803–814.

Reviews and syntheses: Parameter identification in marine planktonic ecosystem modelling. M. Schartau, P. Wallhead, J. Hemmings, U. Löptien, I. Kriest, S. Krishna, B.A. Ward, T. Slawig and A. Oschlies (2017). Biogeosciences, 14 (6) 1647-1701.

Marine mixotrophy increases trophic transfer efficiency, mean organism size, and vertical carbon flux. B.A. Ward and M.J. Follows (2016). Proceedings of the National Academy of Sciences, USA, 113 (11) 2958-2963.



Temperature-correlated changes in phytoplankton community structure are restricted to polar waters. B.A. Ward (2016). PloS one, 10 (8) e0135581.

When everything is not everywhere but species evolve: an alternative method to model adaptive properties of marine ecosystems. B. Sauterey, B.A. Ward, M.J. Follows, C. Bowler and D. Claessen (2015). Journal of Plankton Research, 37 (1), 28-47. [Featured Article.]


Modelling spatial and temporal patterns in size-structured marine plankton communities: top–down and bottom–up controls. B.A. Ward, S. Dutkiewicz and M.J. Follows (2014). Journal of Plankton Research, 37 (1), 28-47. [Featured Article.]

The impact of fine-scale turbulence on phytoplankton community structure. A.D. Barton, B.A. Ward, R.G. Williams and M.J. Follows (2014). Limnology and Oceanography: Fluids and Environments, 4 (1), 34-49.

Maximal foraging with active prey-switching: a multi-species functional response. S.M. Vallina, B.A. Ward, S. Dutkiewicz and M.J. Follows (2014). Progress in Oceanography, 120 (1), 93-109.

Understanding predicted shifts in diazotroph biogeography using resource competition theory. S. Dutkiewicz, B.A. Ward, J.R. Scott and M.J. Follows (2014). Biogeosciences, 11, 5445-5461.


Iron, phosphorus and nitrogen supply ratios define the biogeography of nitrogen fixation. B.A. Ward, S. Dutkiewicz, C.M. Moore, and M.J. Follows (2013). Limnology and Oceanography, 58 (6), 2059–2075.

When is a biogeochemical model too complex? Objective model reduction for North Atlantic time-series sites. B.A. Ward, M. Schartau, A. Oschlies, A.P. Martin, M.J. Follows and T.R. Anderson (2013). Progress in Oceanography, 116, 49-65.

The biogeography of marine plankton traits. A.D. Barton, A.J. Pershing, E. Litchman, N.R. Record, K.F. Edwards, Z.V. Finkel, T. Kiørboe, B.A. Ward (2013). Ecology Letters, 16 (4), 522-534.

On the roles of cell size and trophic strategy in North Atlantic diatom and dinoflagellate communities. A.D. Barton, Z.V. Finkel, B.A. Ward, D.G. Johns and M.J. Follows (2013). Limnology and Oceanography, 58 (1), 254-266.


A size structured food-web model for the global ocean. B.A. Ward, S. Dutkiewicz, O. Jahn and M.J. Follows (2012). Limnology and Oceanography, 57 (6), 1877-1891.

Interconnection of nitrogen fixers and iron in the Pacific Ocean: Theory and numerical simulations. S. Dutkiewicz, B.A. Ward, F. Monteiro and MJ Follows (2012). Global Biogeochemical Cycles, 26, GB1012.

Multi-decadal variability and trends of the temperature of the northwest European continental shelf: a model-data synthesis. J. Holt, S. Hughes, J. Hopkins, S.L. Wakelin, N.P. Holliday, S. Dye, C. González-Pola, S.S. Hjøllo, K.A. Mork, G. Nolan, R. Proctor, J. Read, T. Shammon, T. Sherwin, T. Smyth, G. Tattersall, B.A. Ward and K.H. Wiltshire (2012). Progress in Oceanography, 106, 96-117


Biophysical aspects of resource acquisition and competition in algal mixotrophs. B.A. Ward, S. Dutkiewicz, A.D. Barton and M.J. Follows (2011). The American Naturalist, 178 (1), 98-112.


Parameter optimisation techniques and the problem of underdetermination in marine biogeochemical models. B.A. Ward, M.A.M. Friedrichs, T.R. Anderson, A. Oschlies (2010). Journal of Marine Systems, 81 (1-2), 34-43.

Parameter optimization and uncertainty analysis in a model of oceanic CO2 uptake using a hybrid algorithm and algorithmic differentiation. J. Rückelt, V. Sauerland, T. Slawig, A. Srivastav, B.A. Ward, C Patvardhan (2010). Nonlinear Analysis: Real World Applications, 11, 3993-4009.


Phytoplankton growth conditions in the Irminger Sea, North Atlantic. B.A. Ward and J.J. Waniek (2007). Marine Ecology Progress Series, 334, 47-61.